Cardiff University, Fujian Medical University, Fujian Provincial Center for Disease Control and Prevention, Fujian Medical University, Fujian Medical University, Fujian Medical University, Fujian Medical University, Chinese Academy of Medical Sciences & Peking Union Medical College (CAMS & PUMC), Chinese Academy of Medical Sciences - NHC Key Laboratory of Systems Biology of Pathogens, Fujian Medical University, Fujian Medical University, Cardiff University, Fujian Medical University - Department of Nursing and Fujian Medical University - Fujian Provincial Hospital