MRC London Institute of Medical Sciences, MRC London Institute of Medical Sciences, MRC London Institute of Medical Sciences, University of Edinburgh - Institute of Evolutionary Biology, California Institute of Technology - Division of Biology and Biological Engineering, California Institute of Technology - Division of Biology and Biological Engineering, New York University (NYU) - Department of Biology, New York University (NYU) - Department of Biology, New York University (NYU) - Department of Biology, MRC London Institute of Medical Sciences, University of Edinburgh - Institute of Evolutionary Biology and MRC London Institute of Medical Sciences