Seoul National University - Department of Immunology and Molecular Microbiology, Seoul National University - Department of Immunology and Molecular Microbiology, Seoul National University - Department of Immunology and Molecular Microbiology, Seoul National University - Department of Immunology and Molecular Microbiology, Seoul National University - Oral Medicine and Oral Diagnosis, Government of the United States of America - Experimental Immunology Branch, Seoul National University - Department of Immunology and Molecular Microbiology, Seoul National University - Department of Immunology and Molecular Microbiology, Catholic University of Korea - Division of Rheumatology, Catholic University of Korea - Division of Rheumatology and Seoul National University - Department of Immunology and Molecular Microbiology