University of Science and Technology of China (USTC), School of Life Sciences, Division of Life Sciences and Medicine, Hefei National Laboratory for Physical Sciences at the Microscale, University of Science and Technology of China (USTC), School of Life Sciences, Division of Life Sciences and Medicine, Hefei National Laboratory for Physical Sciences at the Microscale, University of Science and Technology of China (USTC), School of Life Sciences, Division of Life Sciences and Medicine, Hefei National Laboratory for Physical Sciences at the Microscale, Chinese Academy of Sciences (CAS) - Center for Excellence in Brain Science and Intelligence Technology, Chinese Academy of Sciences (CAS) - Center for Excellence in Brain Science and Intelligence Technology, University of Science and Technology of China (USTC), School of Life Sciences, Division of Life Sciences and Medicine, Hefei National Laboratory for Physical Sciences at the Microscale, University of Science and Technology of China (USTC), School of Life Sciences, Division of Life Sciences and Medicine, Hefei National Laboratory for Physical Sciences at the Microscale and University of Science and Technology of China (USTC) - Hefei National Laboratory for Physical Sciences at the Microscale