Exploratory Research Center on Life and Living Systems (ExCELLS), Rikkyo University, College of Science, Department of Life Science, Toyohashi University of Technology, Department of Applied Chemistry and Life Science, Yamagata University - Department of Food, Life, and Environmental Sciences, RIKEN Center for Sustainable Resource Science, RIKEN Center for Sustainable Resource Science, Hokkaido University - Department of Biological Science and Exploratory Research Center on Life and Living Systems (ExCELLS)