University of Tokyo - Department of Cell Biology and Anatomy, University of Tokyo - Department of Cell Biology and Anatomy, University of Tokyo - Department of Cell Biology and Anatomy, University of Tokyo - Department of Cell Biology and Anatomy, University of Tokyo - Department of Cell Biology and Anatomy, Fukushima Medical University - Department of Neuropsychiatry, Niigata University - Department of Pathology, RIKEN Center for Brain Science - Laboratory for Molecular Psychiatry, Fukushima Medical University - Department of Neuropsychiatry, RIKEN Center for Brain Science - Laboratory for Molecular Psychiatry, University of Tokyo - Department of Cell Biology and Anatomy and University of Tokyo - Department of Cell Biology and Anatomy