State Key Laboratory of Remote Sensing Science, Center for Global Change and Public Health, College of Life Sciences, Beijing Normal University, Department of Microbiology and Immunology, Rega Institute, Laboratory of Evolutionary and Computational Virology, Princeton University, University of Oxford - Department of Zoology, Yunnan Institute of Endemic Diseases Control and Prevention, London School of Hygiene & Tropical Medicine - Centre for Mathematical Modelling of Infectious Diseases and Beijing Normal University (BNU) - State Key Laboratory of Remote Sensing Science