Stanford University - Division of Immunology and Rheumatology, Stanford University - Division of Immunology and Rheumatology, Chinese Academy of Sciences (CAS) - Center for Excellence in Animal Evolution and Genetics, Columbia University Irving Medical Center - Department of Pathology and Cell Biology, Stanford University School of Medicine - Program in Epithelial Biology and Department of Dermatology, Stanford University School of Medicine - Program in Epithelial Biology and Department of Dermatology, Chinese Academy of Sciences (CAS) - Center for Excellence in Animal Evolution and Genetics, Stanford University - Division of Immunology and Rheumatology, Stanford University - Division of Immunology and Rheumatology, Stanford University - Division of Immunology and Rheumatology, Stanford University - Division of Immunology and Rheumatology, Columbia University - Department of Microbiology and Immunology, Stanford University - Division of Immunology and Rheumatology and Stanford University - Division of Immunology and Rheumatology