Harvard University - Department of Genetics, Harvard University, Stanford University - Department of Biology, University of Glasgow, Stanford University, Harvard University - Department of Genetics, Harvard University - Drosophila RNAi Screening Center, Harvard University - Drosophila RNAi Screening Center, Stanford University, Stanford University - Department of Bioengineering, Stanford University - Department of Bioengineering, Stanford University - Department of Biology, University of Southern California - Department of Stem Cell Biology and Regenerative Medicine, University of Glasgow and Harvard University - Department of Genetics