Shandong University, Terrestrial Wildlife Rescue and Epidemic Diseases Surveillance Center of Guangxi, Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology - State Key Laboratory of Pathogen and Biosecurity, Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, Chinese Academy of Sciences (CAS) - CAS Key Laboratory of Pathogenic Microbiology and Immunology, Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, Terrestrial Wildlife Rescue and Epidemic Diseases Surveillance Center of Guangxi, Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, Shandong University - Institute of EcoHealth, Terrestrial Wildlife Rescue and Epidemic Diseases Surveillance Center of Guangxi, Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, Terrestrial Wildlife Rescue and Epidemic Diseases Surveillance Center of Guangxi, Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, Terrestrial Wildlife Rescue and Epidemic Diseases Surveillance Center of Guangxi, Terrestrial Wildlife Rescue and Epidemic Diseases Surveillance Center of Guangxi, Terrestrial Wildlife Rescue and Epidemic Diseases Surveillance Center of Guangxi, Terrestrial Wildlife Rescue and Epidemic Diseases Surveillance Center of Guangxi, Terrestrial Wildlife Rescue and Epidemic Diseases Surveillance Center of Guangxi, Guangxi Medical University - Guangxi Collaborative Innovation Center for Biomedicine, Life Sciences Institute, Shandong University - Institute of EcoHealth and Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity