St Mary’s Campus, Imperial College - Department of Infectious Disease Epidemiology, University College London - Department of Genetics, Evolution and Environment, St Mary’s Campus, Imperial College - Department of Infectious Disease Epidemiology, St Mary’s Campus, Imperial College - Department of Infectious Disease Epidemiology, University of Copenhagen - Section for Evolutionary Genomics, University of Dar es Salaam - Department of Zoology and Wildlife Conservation, Manchester Metropolitan University - Department of Natural Sciences, Wild Planet Trust, Paignton Zoo - Field Conservation and Research Department, Manchester Metropolitan University - Department of Natural Sciences, Zoological Society of London - Institute of Zoology, Zoological Society of London - Institute of Zoology, University of Copenhagen - Section for Evolutionary Genomics, North-West University - Unit for Environmental Sciences and Management and St Mary’s Campus, Imperial College - Department of Infectious Disease Epidemiology