Chinese Academy of Sciences (CAS) - Key Laboratory of Animal Ecology and Conservation Biology, Beijing Forestry University - School of Ecology and Nature Conservation, Chinese Academy of Sciences (CAS) - Key Laboratory of Animal Ecology and Conservation Biology, Chinese Academy of Sciences (CAS) - CAS Key Laboratory of Tropical Marine Bio-resources and Ecology, Chinese Academy of Fishery Sciences - Key Laboratory of Recreational Fisheries, Ministry of Agriculture and Rural Affairs, Tibet Agriculture & Animal Husbandry University - National Forest Ecosystem Observation & Research Station of Nyingchi Tibet, Tibet Plateau Institute of Biology, Chinese Academy of Sciences (CAS) - Key Laboratory of Animal Ecology and Conservation Biology, Chinese Academy of Sciences (CAS) - Key Laboratory of Animal Ecology and Conservation Biology, Beijing Forestry University - School of Landscape Architecture, Beijing Forestry University - College of Biological Sciences and Biotechnology, Beijing Forestry University - College of Biological Sciences and Biotechnology, IUCN China country office, IUCN China country office, Chinese Academy of Sciences (CAS), Hanjiang Normal University - Institute of Chemistry and Environmental Engineering and Chinese Academy of Sciences (CAS) - Key Laboratory of Animal Ecology and Conservation Biology