Institute of Applied Ecology - Huitong Experimental Station of Forest Ecology, Institute of Applied Ecology - Huitong Experimental Station of Forest Ecology, Institute of Applied Ecology - Huitong Experimental Station of Forest Ecology, Institute of Applied Ecology - Huitong Experimental Station of Forest Ecology, Institute of Applied Ecology - Huitong Experimental Station of Forest Ecology, Institute of Applied Ecology - Huitong Experimental Station of Forest Ecology, Chinese Academy of Sciences (CAS) - Key Laboratory of Forest Ecology and Management, Hunan Academy of Forestry, Liaoning Academy of Agricultural Sciences, affiliation not provided to SSRN, Institute of Applied Ecology - Huitong Experimental Station of Forest Ecology and Institute of Applied Ecology - Huitong Experimental Station of Forest Ecology