Nagasaki University - School of Tropical Medicine and Global Health, MAIKEN, Saitama Medical University - Department of Microbiology, Saitama Medical University - Department of Infectious Disease and Infection Control, Nagasaki University - Graduate School of Tropical Medicine and Global Health, London School of Hygiene & Tropical Medicine - Department of Clinical Research, Nagasaki University - Department of Nursing Sciences and Nagasaki University - Department of Immunogenetics