University College London - UCL Division of Surgery and Interventional Science, Wellcome Sanger Institute - Wellcome Genome Campus, University College London - UCL Institute of Immunity and Transplantation, University College London - Department of Histopathology, University College London - Department of Histopathology, Guy's and St Thomas' NHS Foundation Trust, University College London - UCL Division of Surgery and Interventional Science, Wellcome Sanger Institute, University College London - UCL Institute of Immunity and Transplantation, University College London - UCL Division of Surgery and Interventional Science and Wellcome Sanger Institute - Wellcome Genome Campus