Beijing Normal University, Beijing Key Laboratory of Gene Resource and Molecular Development, Beijing Normal University, Beijing Key Laboratory of Gene Resource and Molecular Development, Beijing Normal University, Beijing Key Laboratory of Gene Resource and Molecular Development, Beijing Normal University, Beijing Key Laboratory of Gene Resource and Molecular Development, Beijing Normal University, Beijing Key Laboratory of Gene Resource and Molecular Development, Beijing Normal University, Beijing Key Laboratory of Gene Resource and Molecular Development and Beijing Normal University, Beijing Key Laboratory of Gene Resource and Molecular Development