Fudan University - State Key Laboratory of Medical Neurobiology and MOE Frontiers Center for Brain Science, Fudan University - State Key Laboratory of Medical Neurobiology and MOE Frontiers Center for Brain Science, Fudan University - State Key Laboratory of Medical Neurobiology and MOE Frontiers Center for Brain Science, Fudan University - State Key Laboratory of Medical Neurobiology and MOE Frontiers Center for Brain Science, Fudan University - State Key Laboratory of Medical Neurobiology and MOE Frontiers Center for Brain Science, Fudan University - Key Laboratory of Medical Molecular Virology (MOE/NHC/CAMS), ShanghaiTech University - School of Life Science and Technology, Shanghai University of Traditional Chinese Medicine - School of Rehabilitation Science, Fudan University - State Key Laboratory of Medical Neurobiology and MOE Frontiers Center for Brain Science, Fudan University - State Key Laboratory of Medical Neurobiology and MOE Frontiers Center for Brain Science, Fudan University - State Key Laboratory of Medical Neurobiology and MOE Frontiers Center for Brain Science, Southern University of Science and Technology - Shenzhen Key Laboratory of Smart Healthcare Engineering, Southern University of Science and Technology - Shenzhen Key Laboratory of Smart Healthcare Engineering, Fudan University - Key Laboratory of Medical Molecular Virology (MOE/NHC/CAMS), ShanghaiTech University - School of Life Science and Technology and Fudan University - State Key Laboratory of Medical Neurobiology and MOE Frontiers Center for Brain Science