South China Agricultural University, South China Agricultural University, Guangdong Academy of Agricultural Sciences, South China Agricultural University, South China Agricultural University, South China Agricultural University, Guangdong Academy of Agricultural Sciences, Guangdong Academy of Agricultural Sciences, Guangdong Academy of Agricultural Sciences, Guangdong Academy of Agricultural Sciences, Guangdong Academy of Agricultural Sciences, National Avian Influenza Para-Reference Laboratory, Guangdong Academy of Agricultural Sciences and South China Agricultural University