affiliation not provided to SSRN, Universidade Federal do Rio de Janeiro (UFRJ), Universidade Federal do Rio de Janeiro (UFRJ), affiliation not provided to SSRN, Federal University of Para, Federal University of Para, Emilio Goeldi Museum of Para, Smithsonian Tropical Research Institute, affiliation not provided to SSRN, affiliation not provided to SSRN, Rio de Janeiro State University, Rio de Janeiro State University, Federal University of Cariri, affiliation not provided to SSRN, affiliation not provided to SSRN and Federal University of Para