Central South University of Forestry and Technology, Central South University of Forestry and Technology, Central South University of Forestry and Technology, Central South University of Forestry and Technology, affiliation not provided to SSRN, Central South University of Forestry and Technology, Central South University of Forestry and Technology, Central South University of Forestry and Technology, Central South University of Forestry and Technology, Chinese Academy of Sciences (CAS) - South China Botanical Garden, Central South University of Forestry and Technology and Central South University of Forestry and Technology