Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, Chinese Academy of Sciences (CAS) - Guangdong Key Laboratory of Nanomedicine, Gannan Medical University, Beijing Institute of Microbiology and Epidemiology - State Key Laboratory of Pathogen and Biosecurity, Shandong University, Shandong University - Institute of EcoHealth, Shandong University - Institute of EcoHealth, Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology - State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology - State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology - State Key Laboratory of Pathogen and Biosecurity, Beijing Institute of Microbiology and Epidemiology - State Key Laboratory of Pathogen and Biosecurity, Chinese PLA Center for Disease Control and Prevention, Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, ManZhouLi Customs, ManZhouLi Customs, ManZhouLi Customs, Beijing Institute of Microbiology and Epidemiology - State Key Laboratory of Pathogen and Biosecurity, Chinese Academy of Sciences (CAS) - Guangdong Key Laboratory of Nanomedicine, Chinese Academy of Sciences (CAS) - Guangdong Key Laboratory of Nanomedicine, Academy of Military Sciences - State Key Laboratory of Pathogen and Biosecurity, Shandong University - Institute of EcoHealth and Independent