University of Oxford - Nuffield Department of Medicine, University of Oxford - Department of Zoology, University of the Free State, University of Oxford, University of Oxford - Nuffield Department of Medicine, Cardiff and Vale University Health Board - Public Health Wales Microbiology Cardiff, Stellenbosch University, Government of Uganda - Uganda Virus Research Institute (UVRI), University of the Free State, University of Oxford - Department of Zoology, MRC/UVRI and LSHTM Research Unit, MRC/UVRI and LSHTM Uganda Research Unit, Makerere University and University of Oxford - Nuffield Department of Medicine