Shiga University of Medical Science, Shiga University of Medical Science, Tokyo Metropolitan Institute of Medical Science, Shiga University of Medical Science, Shiga University of Medical Science, Tokyo Metropolitan Institute of Medical Science, Tokyo Metropolitan institute of Medical Science - Department of Microbiology and Cell Biology, Tokyo Metropolitan Institute of Medical Science, Tokyo Metropolitan Institute of Medical Science, Tokyo Metropolitan Institute of Medical Science, National Institute of Infectious Diseases, affiliation not provided to SSRN, affiliation not provided to SSRN, Tokyo Metropolitan institute of Medical Science - Department of Microbiology and Cell Biology and Shiga University of Medical Science