Nanjing Forestry University, affiliation not provided to SSRN, Nanjing Medical University - Department of Otorhinolaryngology-Head and Neck Surgery, Nanjing Normal University, Nanjing Forestry University, Nanjing Normal University, Jilin University (JLU) - Key Laboratory of Zoonosis Research, Nanjing Forestry University and Nanjing Normal University