Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, affiliation not provided to SSRN, Zhongkai University of Agriculture and Engineering, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Zhongkai University of Agriculture and Engineering and Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute