affiliation not provided to SSRN, affiliation not provided to SSRN, affiliation not provided to SSRN, affiliation not provided to SSRN, INRA, affiliation not provided to SSRN, affiliation not provided to SSRN, affiliation not provided to SSRN, Sidi Mohamed Ben Abdellah University - Laboratory of Functional Ecology and Environment, Sidi Mohamed Ben Abdellah University - Laboratory of Functional Ecology and Environment, affiliation not provided to SSRN, Sidi Mohamed Ben Abdellah University - Laboratory of Functional Ecology and Environment, affiliation not provided to SSRN, affiliation not provided to SSRN, affiliation not provided to SSRN, affiliation not provided to SSRN, Sidi Mohamed Ben Abdellah University - Laboratory of Functional Ecology and Environment, Sidi Mohamed Ben Abdellah University - Laboratory of Functional Ecology and Environment, Université de Toulouse, affiliation not provided to SSRN and affiliation not provided to SSRN