Kobe University, Ehime University, Kobe University, Kobe University, Kyoto University, Hokkaido University, Tokyo University of Agriculture, Saitama University, National Institute for Basic Biology (NIBB) - Division of Cellular Dynamics, The University of Osaka, National Institute for Basic Biology (NIBB) - Division of Cellular Dynamics, The University of Osaka, Kobe University, Kobe University - Graduate School of Science and Independent