Hokkaido University, University of Missouri, National Trust for Nature Conservation, Hokkaido University - Research Center for Zoonosis Control, Hokkaido University, affiliation not provided to SSRN, National Trust for Nature Conservation, affiliation not provided to SSRN, Hokkaido University, affiliation not provided to SSRN, affiliation not provided to SSRN, affiliation not provided to SSRN, Long Island University, National Trust for Nature Conservation and Hokkaido University