Fujian Normal University, Fujian Normal University, Fujian Normal University, Fujian Normal University, Chinese Academy of Sciences (CAS) - Yantai Institute of Coastal Zone Research, CSIC, Global Ecology Unit CREAF-CSIC-UAB, Fujian Normal University, Fujian Normal University, Fujian Normal University, Fujian Normal University, Fujian Normal University, Fujian Normal University and Autonomous University of Barcelona