Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Guangdong University of Technology, affiliation not provided to SSRN, affiliation not provided to SSRN, Yamagata University, Hainan University, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute and Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute