Vanderbilt University - Department of Cell and Developmental Biology, Vanderbilt University - Department of Biomedical Engineering, Vanderbilt University - Department of Cell and Developmental Biology, Vanderbilt University - Department of Biomedical Engineering, Vanderbilt University - Department of Cell and Developmental Biology, University of Michigan at Ann Arbor - Department of Cell and Developmental Biology, Vanderbilt University - Department of Cell and Developmental Biology, Vanderbilt University - Department of Biomedical Engineering and Vanderbilt University - Department of Cell and Developmental Biology