Fudan University - State Key Laboratory of Genetic Engineering, Shanghai Academy of Chinese Medical Sciences, Fudan University - Department of Biochemistry and Molecular Biology, Fudan University - State Key Laboratory of Genetic Engineering, Fudan University - Ministry of Education Key Laboratory of Contemporary Anthropology, Nantong University - Taizhou People's Hospital, Shanghai Academy of Chinese Medical Sciences, Fudan University - State Key Laboratory of Genetic Engineering, Fudan University - Department of Biochemistry and Molecular Biology, University of Texas at Houston, University of Texas at Houston, Fudan University - Collaborative Innovation Center for Genetics and Development, Fudan University, Fudan University - Department of Biochemistry and Molecular Biology and Fudan University