University of Tokyo - Laboratory of Neuropathology and Neuroscience, University of Tokyo - Laboratory of Neuropathology and Neuroscience, University of Tokyo - Laboratory of Neuropathology and Neuroscience, University of Tokyo - Laboratory of Neuropathology and Neuroscience, Tokyo Medical and Dental University - Department of Lipid Biology, RIKEN Center for Brain Science - Laboratory for Proteolytic Neuroscience, RIKEN Center for Brain Science - Laboratory for Proteolytic Neuroscience, Mayo Clinic - Florida - Department of Neuroscience, Tohoku University - Department of Experimental Immunology, Tokyo Medical and Dental University - Department of Lipid Biology and University of Tokyo - Laboratory of Neuropathology and Neuroscience