RIKEN Center for Brain Science, University of Tokyo - Laboratory of Neuropathology and Neuroscience, University of Tokyo - Laboratory of Neuropathology and Neuroscience, University of Tokyo - Laboratory of Neuropathology and Neuroscience, University of Tokyo - Laboratory of Neuropathology and Neuroscience, Tokyo Medical and Dental University, RIKEN Center for Brain Science, Mayo Clinic - Florida - Department of Neuroscience, Tohoku University, Tokyo Medical and Dental University and University of Tokyo - Laboratory of Neuropathology and Neuroscience