Guizhou University, Guizhou University, Sichuan Academy of Grassland Sciences, Sichuan Academy of Grassland Sciences, Sichuan Academy of Grassland Sciences, Sichuan Academy of Grassland Sciences, Guizhou University, Southwest Minzu University, Qinghai University, Qinghai University, Guizhou University, Tibet Academy of Agricultural and Animal Husbandry Sciences, Tibet Academy of Agricultural and Animal Husbandry Sciences, Tibet Institute of Modern Life and Health, Tibet Jingliang Agriculture and Animal Husbandry Industry Development Co., Guizhou University and Sichuan Academy of Grassland Sciences