Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, South China Agricultural University, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Ocean University of China - Sanya Tropical Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, South China University of Technology and Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute