affiliation not provided to SSRN, Cairo University, affiliation not provided to SSRN, Cairo University, affiliation not provided to SSRN, affiliation not provided to SSRN, South China Agricultural University, Guangdong Academy of Agricultural Sciences - Guangdong Key Laboratory for Crop Germplasm Resources Preservation and Utilization, Guangdong Academy of Agricultural Sciences, Guangdong Academy of Agricultural Sciences and affiliation not provided to SSRN