affiliation not provided to SSRN, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences - South China Sea Fisheries Research Institute, Shanghai Ocean University, Zhongkai University of Agriculture and Engineering and Ocean University of China - Sanya Tropical Fisheries Research Institute